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The best way to get around Tuscany is by car - this gives you the freedom to explore all the towns nearby and yummy restaurants in the country. You'll want to get out and explore the region so you can really get a feel for it and appreciate its charm and beauty. If you can't drive a stick shift, make sure to request an automatic because most Italians drive stick. If you think gas is expensive in the U.S., just wait til you get a load of the prices in Europe - you may think it's cheap until you realize it's being sold by the liter, not the gallon.
There are many toll and free highways - and the road conditions are generally pretty good, so you shouldn't have too much trouble. Driving is on the right - which you might not expect if you've been driving around any other parts of Europe.
There is also public transportation in the form of buses and trains. Buses are usually the better option - there are a number of lines that go between the smaller towns and cities. Sometimes it can be tough for non-native speakers to get information, but it's worth the hassle.
You can bike around Tuscany, too. There are biking tours or you can rent a bike on your own and find routes in and around the city.
Also, boats and ferries are a way to get around the area. If you want to explore the islands from the mainland, this is the only way to get around.